Detectives investigating alleged phone-hacking at the News of the World have arrested the paper's chief reporter and former head of news.
Neville Thurlbeck, 50, and Ian Edmondson, 42, were held by Scotland Yard detectives when they voluntarily attended separate police stations in south west London.
A Metropolitan Police spokesman later said two men who had been arrested in connection with the investigation had been released on police bail to return in September.
Officers questioned the pair on suspicion of conspiring to intercept communications and unlawfully intercepting voicemail messages.
The arrests underlined the Metropolitan Police's determination to investigate the scandal thoroughly after criticism that an earlier inquiry was inadequate.
Thurlbeck is a veteran reporter who has brought in some of the News of the World's most famous scoops, while Edmondson was sacked as the paper's assistant editor (news) in January after evidence emerged linking him to phone-tapping.
They were the first people arrested since Scotland Yard reopened its inquiry into claims that staff at the top-selling Sunday newspaper hacked into the answerphone messages of celebrities, politicians and royals.
A committee of MPs heard allegations in 2009 that a transcript of voicemail messages between Professional Footballers' Association boss Gordon Taylor and his legal adviser was prepared for Thurlbeck.
Edmondson's solicitor, Eddie Parladorio, stressed that his client attended the police station voluntarily and has not been charged.
Scotland Yard has endured repeated criticism over its handling of its original phone-hacking inquiry, which led to the conviction of News of the World royal editor Clive Goodman and private investigator Glenn Mulcaire in 2007.